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Rays I
ver. 2.1
Rays. Kakegawa. Rays I. Rays II. Miliobatis


Cretaceous Paleocene Eocene Miocene

1. Cretaceous


Myledaphus bipartitus Upper Cretaceous, Drumheller, Alberta, Canada. Rhinobatoidei incertae sedis, Genus Myledaphus Cope, 1876


Ischyrhiza avonicola Santonian,Tyk Butak, East Mangyshlak, Western Kazakhstan. Family Sclerorhynchidae Cappetta 1974, Genus Ischyrhiza Leidy, 1856


Ptychotrygon sp. Santonian,Tyk Butak, East Mangyshlak, Western Kazakhstan. Family incertae sedis, Genus Ptychotrygon Jaekel, 1894


Rhinobatos casieri Santonian,Tyk Butak, East Mangyshlak, Western Kazakhstan. Family Rhynchobatidae Garman 1913, Genus Rhinobatos Linck, 1790


Rhombodus microdon Maarstrichtian, Ouled Abdoun Basin, Morocco. Family Rhombodontide Cappetta, Genus Rhombodus Dames, 1881



2. Paleocene


Squatirhina numidicaThanetian, Oueld Zem Basin, Morocco. Rhinobatoidei incertae sedis, Genus Squatirhina Casier, 1947


Dasyatis globidens (Ishaquia globidens) Danian, Oueld Zem Basin, Morocco. Family Dasyatidae Jordan, 1888, Genus Dasyatis Rafinesque, 1810


Dasyatis hexagonalis Danian, Louis Gentil Basin, Morocco. Family Dasyatidae Jordan, 1888, Genus Dasyatis Rafinesque, 1810


Heterotorpedo sp. Thanetian,Oueld Zem Basin, Morocco. Dasyatoidea incertae familiae, Genus Heterotorpedo Ward, 1983



3. Eocene


Hypolophodon cf. sylvestris early Eocene, Nanjemoy Fm. (Fisher/Sullivan Site), Stafford County, Virginia. Family Dasyatidae Jordan, 1888, Genus Hypolophodon Cappetta, 1980


Rhynchobatus sp.(Wedgefish shark) Lakeniaan, Lede Fm., Oosterzele Quarry, Belgium. Family Rhynchobatidae Garman, 1913, Genus Rhynchobatus Muller & Henle 1837


Jacquhermania duponti Egem Sands, Egem village, Belgium. Family Gymnuridae Fowler, 1934, Genus Jacquhermania Cappetta, 1982


Coupatezia woutersi Fisher-Sullivan Bone Bed(Muddy Creek), Nanjemoy Fm., Potapaco Member, Stafford County, Virginia. Dasyatoidea incertae familiae, Genus Coupatezia Cappetta, 1982


Aetobatus sp. Middle Eocene, Cook Mountain FormationPerkins, Louisiana. Family Myliobatidae Bonaparte, 1838, Genus Aetobatus Blainville, 1816


Genus Myliobatis Cuvier, 1817


Family Rhinopteridae Jordan & Everman, 1896, Genus Rhinoptera Cuvier, 1829


Rhinobatos bruxelliensis (Guitar fish) Eocene, Egem Sands, Egem village, Belgium. Genus Rhinobatos Linck, 1790


` melenhorsti Family Mobulidae Gill 1893, Genus Archaeomanta Herman, 1979


Burnhamia daviesi Ypresian, Genus Burnhamia Cappetta, 1976


Burnhamia woodwardi Eocene, Bartonian, Tusbair Mangyshlak, W. Kazakhstan, Genus Burnhamia Cappetta, 1976



4. Miocene

Pungo River Formation
Beaufort County, North Carolina


Raja dux Family Rajidae Bonaparte 1831 Genus Raja Linnaeus, 1758


Dasyatis cavernosa GenusDasyatis Rafinesque, 1810


Manta fragilis Family Mobulidae Gill, 1893 GenusManta Bancroft, 1829


Mobula pectinata GenusMobula Rafinesque, 1810


Mobula loupianensis GenusMobula Rafinesque, 1810


Plinthicus stenodon GenusPlinthicus Cope, 1869L




20090713 Fumio Nakagawa

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